All vowels are to be pronounced with voice (ghosha) and strength (bala), so that the Udgâtri may give strength to Indra. All sibilants are to be pronounced, neither as if swallowed (agrasta), nor as if thrown out (nirasta), but well opened (vivrita), so that the Udgâtri may give himself to Pragâpati. All consonants are to be pronounced slowly, and without crowding them together, so that the Udgâtri may withdraw himself from Mrityu.

In plain words: This explains how to pronounce the sounds. Vowels should be sung loud and strong to give strength to Indra. Hissing sounds should be clear and open, not swallowed or spat out. Consonants should be sung slowly, without running them together, so the singer keeps himself away from Death.